Iwata, Eri, Department of Science and Technology, Iwaki Meisei University, 5-5-1 Chuoudai, Ihino, Iwaki, Fukushima 970-8551, Japan; Faculty of Veterinary Medicine, Okayama University of Science, 1-3, Ikoinooka, Imabari City, Ehime, 794-8555, Japan; Institute of Osaka Marine Research, 19-34, Shimodachou, Nishi Ku, Sakai City, Osaka, 593-8329, Japan, Japan